Dense halophilous grasslands and scrubs occupying the estuarine flats generated by the retention of suspended particles (by means of a self-reinforcing successional process) during periods of inundation in high-tide. When mature and sufficiently elevated, they are at most briefly submersed only during spring high tides.
Protection
Those stands rich in shrubs like Sarcocornia pruinosa and Suaeda vera in sandy soils (see below) are referable to the code 1420 ("Mediterranean and thermo-Atlantic halophilous scrubs [Sarcocornetea fruticosae]") in the EU Habitat Directive.
Floristic Composition
Floristic Composition
Frequent species show the percentage of occurrence of species in each habitat, with higher values indicating more observations within the total number of samples. Indicator species show the fidelity of species to each habitat, with higher values indicating a stronger association, based on the IndVal metric (Dufrêne & Legendre, 1997). All species values were obtained from numerical classification (Jiménez-Alfaro et al. 2025). The classification of anthropogenic habitats (EUNIS code “V”) was conducted separately (Fernández-Pascual et al. 2025).
References
Dufrêne, M., & Legendre, P. (1997). Species assemblages and indicator species: the need for a flexible asymmetrical approach. Ecological monographs, 67(3), 345-366.
Fernández-Pascual, E., González-García, V., Ivesdal, G., Lázaro-Lobo, A., & Jiménez-Alfaro, B. (2024). Classification and Characterization of Anthropogenic Plant Communities in the Northwestern Iberian Peninsula. Applied Vegetation Science, 28(1), e70010. https://doi.org/10.1111/avsc.70010
Jiménez-Alfaro, B., González Le Barbier J., Carlón L., Álvarez Martínez JM, Espinosa del Alba, C., Fernández-Pascual, E., González-García V., Lázaro-Lobo A, Mendieta-Leiva, Roces Díaz JV (2025) Regionalization of European EUNIS habitat classification for effective ecosystem assessment. In prep.
List of phytosociological associations described in the Ibero-Atlantic region for each habitat. The association concepts mainly follow Rivas-Martinez et al. (2001), with further updates by Fernández-Prieto et al. (2023), and others.
References
Rivas-Martinez, S., Fernández González, F., Loidi Arregui, J., Fernández Lousã, M. & Penas Merino, A. (2001) Sintaxonomical checklist of vascular plant communities of Spain and Portugal to association level. Itinera Geobotanica 14., 5-341.
Fernández Prieto, J.A., Amigo, J., Bueno, A., Herrera, M., Rodríguez-Guitián, M.A. & Loidi, J. (2023) Bosques y orlas forestales de los territorios atlánticos del Noroeste Ibérico. Guineana, 23, 3–240 https://doi.org/10.1387/guineana.25055
Agrostio stoloniferae-Juncetum maritimi
Agrostio-Paspaletum vaginati
Inulo crithmoidis-Elytrigietum athericae
Junco maritimi-Caricetum extensae
Junco maritimi-Phragmitetum australis
Puccinellio maritimae-Juncetum maritimi
Scirpetum compacti
Scirpetum compacto-tabernaemontani
Spergulario marinae-Cotuletum coronopifoliae
Relations with other habitat types
These communities replace MA224 as a consequence of sediment retention. Local deposits of litter harbor. tpically around the high-tide driftline, nitrophilous communities recalling those of the EUNIS unit MA221; whereas, in mown and pastured areas, meadows dominated by Agrostis stolonifera are common. Very locally and episodically, in ephemeral muddy clearings and margins of brackish ponds, beds of Eleocharis parvula like those typified by EUNIS as MA228 develop. Similarly, in areas almost never bathed by muddy waters, sediments are mostly sandy (sand dries-up more easily and can be mobilised by wind and deposited above the spring high-tide line) and halophytic shrubs prevail (these communities would be referable to EUNIS MA222 if further data enable their statistical recognition). Lastly, this unit also includes tall helophytic beds of Cyperaceae, especially Bolboschoenus maritimus, fringing the main channel in inner areas of the estuaries, mainly in those created by large rivers where the contribution of freshwater is higher, communities whose best match in EUNIS, in case of an eventual statistical recognition, would be Q517 (see Q51x).
This unit is better or worse represented in almost all estuarine systems along the Iberoatlantic shores. Their extension and internal diversity, however, has been particularly reduced, especially in the last two centuries, by processes of polderization to gain agricultural, urban or industrial land.
Regional distribution
Occurrence data show the locations of vegetation plots from SIVIM (Font, 2023) classified into habitat types by Jiménez-Alfaro et al. (2025). Distribution errors (outliers) may occur in vegetation plots with few species or transitional ecological conditions. The potential area of occupancy shows the environmental suitability predicted for each habitat (González Le Barbier et al. 2025), with higher values indicating more suitable climate and soil for the habitat.
References
Font, X. (2023). Sistema de Información de la vegetación Ibérica y Macaronésica. Version 1.6. Banc de dades de biodiversitat de Catalunya. http://www.sivim.info/sivi/.
Jiménez-Alfaro, B., González Le Barbier J., Carlón, L., Espinosa del Alba, C., Fernández-Pascual, E., González-García, V., Lázaro-Lobo, A. Adapting EUNIS habitat classification at the regional scale: a case study in the Ibero-Atlantic biogeographic region. In prep.
González Le Barbier, J., Ubaldi, T., Lenoir, J., Roces Díaz, J. & Jiménez-Alfaro, B. (2025) Modelling the area of occupancy of habitat types at different successional stages. In prep.
Known occurrences and potential area of occupancy of the habitat type in the study region.